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List 196 - Achatocarpaceae - 2/20/2019


Achatocarpaceae ~ Phytolaccaceae Habit and leaf form. Small trees, or shrubs (the branchlets sometimes spiny). Plants non-succulent. Leptocaul. Leaves alternate; spiral; non-sheathing; simple. Lamina entire; pinnately veined; cross-venulate. Leaves exstipulate. Lamina margins entire. Reproductive type, pollination. Plants dioecious. Female flowers without staminodes. Gynoecium of male flowers absent. Inflorescence, floral, fruit and seed morphology. Flowers aggregated in ‘inflorescences’; in cymes, or in racemes, or in fascicles, or in panicles. The terminal inflorescence unit cymose. Inflorescences small, branched cymes, racemes, panicles or fascicles. Flowers bracteate; small. Free hypanthium absent. Perianth sepaline; 4, or 5; 1 whorled; green. Calyx 4, or 5; 1 whorled; polysepalous; regular; persistent; imbricate. Fruit fleshy; indehiscent; a berry. Dispersal unit the fruit. Fruit 1 seeded. Seeds (strictly) non-endospermic. Perisperm present (copious, mealy). Embryo well differentiated. Cotyledons 2. Embryo curved. Geography, cytology. Sub-tropical to tropical. Warm America. Taxonomy. Subclass Dicotyledonae; Crassinucelli. Dahlgren’s Superorder Caryophylliflorae; Caryophyllales. Cronquist’s Subclass Caryophyllidae; Caryophyllales. APG (1998) Eudicot; core Eudicot; neither Rosid nor Asterid; Caryophyllales. Species 10. Genera 2; Achatocarpus, Phaulothamnus.

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